Physematium

Physematium
	Physematium oreganum
Scientific classification
Kingdom:	Plantae
Clade:	Tracheophytes
Division:	Polypodiophyta
Class:	Polypodiopsida
Order:	Polypodiales
Suborder:	Aspleniineae
Family:	Woodsiaceae
Genus:	Physematium; Kaulf.
Type species
	Physematium molle; Kaulfuss
Species
	See text
Synonyms
	Cheilanthopsis Hieronymus 1920; Hymenocystis Meyer 1831; Protowoodsia Ching 1945; Woodsiopsis Shmakov 2015;

Physematium is a genus of ferns in the family Woodsiaceae. It was treated as a synonym of Woodsia until 2020 when a molecular phylogenetic study showed its distinctiveness.^[1] As of June 2023^[update], Plants of the World Online continued to treat the genus as a synonym of Woodsia.^[2]

Phylogeny

As of June 2023^[update], World Ferns accepted 20 species and two hybrids:^[3]

Phylogeny from Fern Tree of Life^[4]^[5]

Other species include:

Physematium

(Cheilanthopsis)

	P. elongatum (Hooker) Trevis.

	P. indusiosum (Christ) Li Bing Zhang, N.T.Lu & X.F.Gao

	P. kangdingense (H.S.Kung, Li Bing Zhang & X.S.Guo) Li Bing Zhang, N.T.Lu & X.F.Gao

	P. manchuriense (Hook.) Nakai

(Physematium)

P. fragile (Trevis.) Kunze

	P. molle Kaulf.

	P. montevidense (Spreng.) Shmakov

(Woodsiopsis)

P. oreganum (D.C.Eaton) Trevis.

	P. obtusum (Spreng.) Hook.

	P. plummerae (Lemmon) Li Bing Zhang, N.T.Lu & X.F.Gao

	P. cochisense (Windham) Zhang, Lu & Gao

	P. phillipsii (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao

	P. cystopteroides (Windham & Mickel) Zhang, Lu & Gao

	P. neomexicanum (Windham) Li Bing Zhang, N.T.Lu & X.F.Gao

^ Lu, N.T.; Zhou, X.-M.; Zhang, L.; Knapp, R.; Li, C.-X.; Fan, X.-P.; Zhou, L.; Wei, H.-J.; Lu, J.-M., Xu, B.; Peng, Y.-L.; Gao, X.-F. & Zhang, L.-B. (2020), "A global plastid phylogeny of the cliff fern family Woodsiaceae and a two-genus classification of Woodsiaceae with the description of ×Woodsimatium nothogen. nov.", Taxon, 68 (6): 1149–1172, doi:10.1002/tax.12180, S2CID 213706008{{citation}}: CS1 maint: multiple names: authors list (link)
^ "Physematium Kaulf.", Plants of the World Online, Royal Botanic Gardens, Kew, retrieved 2023-06-13
^ Hassler, Michael, "Physematium", World Ferns, retrieved 2023-06-13
^ Nitta, Joel H.; Schuettpelz, Eric; Ramírez-Barahona, Santiago; Iwasaki, Wataru; et al. (2022). "An Open and Continuously Updated Fern Tree of Life". Frontiers in Plant Science. 13: 909768. doi:10.3389/fpls.2022.909768. PMC 9449725. PMID 36092417.
^ "Tree viewer: interactive visualization of FTOL". FTOL v1.7.0 (GenBank release 261). 2024. Retrieved 1 May 2025.

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