The evolutionary ancestry of arthropods dates back to the Cambrian period. The group is generally regarded as monophyletic, and many analyses support the placement of arthropods with cycloneuralians (or their constituent clades) in a superphylum Ecdysozoa. Overall, however, the basal relationships of animals are not yet well resolved. Likewise, the relationships between various arthropod groups are still actively debated. Today, arthropods contribute to the human food supply both directly as food, and more importantly, indirectly as pollinators of crops. Some species are known to spread severe disease to humans, livestock, and crops. (Full article...)
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Five anatomical images of a Drosophila subobscura (male) on a rotting squash, located near Christchurch Park, Ipswich
Drosophila subobscura is a species of fruit fly in the family Drosophilidae. Originally found around the Mediterranean, it has spread to most of Europe and the Near East. It has been introduced into the west coasts of Canada, the United States, and Chile. Its closest relative is Drosophila madeirensis, found in the Madeira Islands, followed by D. guanche, found in the Canary Islands. These three species form the D. subobscura species subgroup. When they mate, males and females perform an elaborate courtship dance, in which the female can either turn away to end the mating ritual, or stick out her proboscis in response to the male's, allowing copulation to proceed. D. subobscura has been regarded as a model organism for its use in evolutionary-biological studies. (Full article...)
Aptostichus barackobamai (also known as the Barack Obama trapdoor spider) is a large species of trapdoor spider in the familyEuctenizidae named after the 44th President of the United States, Barack Obama. The species was first reported by Jason Bond of Auburn University in December 2012 as one of 33 new species of the genusAptostichus. The species is endemic throughout northern California, forming burrows out of silk, dirt, and sand, from which it attacks prey. This spider is identified by its dark red to red-brown coloration with a stripped or chevron patterned abdomen. Breeding occurs during the winter and females tend to be larger than their male counterparts. A. barackobamai is part of the larger Aptostichus icenogleispecies complex, and is a close relative to Aptostichus icenoglei and Aptostichus isabella. (Full article...)
Isopods have various feeding methods: some eat dead or decaying plant and animal matter, others are grazers or filter feeders, a few are predators, and some are internal or external parasites, mostly of fish. Aquatic species mostly live on the seabed or the bottom of freshwater bodies of water, but some taxa can swim for short distance. Terrestrial forms move around by crawling and tend to be found in cool, moist places. Some species are able to roll themselves into a ball as a defense mechanism or to conserve moisture like species in the family Armadillididae, the pillbugs.
There are over 10,000 identified species of isopod worldwide, with around 4,500 species found in marine environments, mostly on the seabed, 500 species in fresh water, and another 5,000 species on land. The order is divided into eleven suborders. The fossil record of isopods dates back to the Carboniferous period (in the Pennsylvanian epoch), at least 300 million years ago, when isopods lived in shallow seas. The name Isopoda is derived from the Greekrootsiso- (from ἴσοςísos, meaning "equal") and -pod (from ποδ-, the stem of πούςpoús, meaning "foot"). (Full article...)
The orderEmbioptera, commonly known as webspinners or footspinners, are a small group of mostly tropical and subtropicalinsects, classified under the subclassPterygota. The order has also been called Embiodea or Embiidina. More than 400 species in 11 families have been described, the oldest known fossils of the group being from the mid-Jurassic. Species are very similar in appearance, having long, flexible bodies, short legs, and only males having wings.
Webspinners are gregarious, living subsocially in galleries of fine silk which they spin from glands on their forelegs. Members of these colonies are often related females and their offspring; adult males do not feed and die soon after mating. Males of some species have wings and are able to disperse, whereas the females remain near where they were hatched. Newly mated females may vacate the colony and find a new one nearby. Others may emerge to search for a new food source to which the galleries can be extended, but in general, the insects rarely venture from their galleries. (Full article...)
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T. tatusia worker from El Campano, Magdalena, Colombia
Tatuidris, or armadillo ant, is a rare genus of ants consisting of a single species, Tatuidris tatusia. The ants are small in size and inhabit the leaf litter of Neotropical forests in Central and South America, from Mexico to Brazil. Workers are ferruginous-colored to dark red and present a distinctive morphology, consisting of a shield-like head with a broad vertex, ventrally-turned heavy mandibles which do not overlap at full closure, and unique among ants – an antenna socket apparatus sitting upside-down. Little is known about the biology of the ants, but they are likely nocturnal and specialist predators.
Tatuidris was first described in 1968 and initially placed in the myrmicinetribe Agroecomyrmecini, together with two fossil genera. Since the original description, the systematic status of the tribe has been the focus of debate. (Full article...)
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Portia schultzi is a species of jumping spider which ranges from South Africa in the south to Kenya in the north, and also is found in West Africa and Madagascar. In this species, which is slightly smaller than some other species of the genusPortia, the bodies of females are 5 to 7 mm (0.20 to 0.28 in) long, while those of males are 4 to 6 mm (0.16 to 0.24 in) long. The carapaces of both sexes are orange-brown with dark brown mottling, and covered with dark brown and whitish hairs lying over the surface. Males have white tufts on their thoraces and a broad white band above the bases of the legs, and these features are less conspicuous in females. Both sexes have tufts of orange to dark orange above the eyes, which are fringed with pale orange hairs. Males' abdomens are yellow-orange to orange-brown with blackish mottling, and on the upper sides are black and light orange hairs, and nine white tufts. Those of females are pale yellow and have black markings with scattered white and orange-brown hairs on the upper side. P. schultzi has relatively longer legs than other Portia, and a "lolloping" gait.
While most jumping spiders focus accurately up to about 75 cm (30 in) away, P. schultzi responds to a maximum of about 10 cm (3.9 in) in good light, and ignores everything in very subdued light. For prey, P. schultzi prefers web-based spiders, then jumping spiders, and finally insects. The females of P. schultzi and other Portia species build "capture webs" to catch prey, and often join their own webs on to web-based spiders to catch the other spiders or their prey.
If a P. schultzi female is mature, a male P. schultzi will try to copulate with her, or cohabit with a subadult female and copulate while she is moulting. They usually mate on a web or on a dragline made by the female, and P. schultzi typically copulates for about 100 seconds, while others in the genus can take several minutes or even several hours. Females try to kill and eat their mates during or after copulation, and subadult females mimic adult females to attract males as prey. Contests between Portia females are violent, and embraces in P. schultzi typically take 20 to 60 seconds. Sometimes, one female knocks the other on her back and the other may be killed and eaten if she does not right herself quickly and run away. When hunting, P. schultzi mature females emit olfactory signals that reduce the risk that any other females, males, or juveniles of the same species may contend for the same prey. (Full article...)
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Myrmecia nigriceps worker from Strangways, Victoria
Myrmecia nigriceps, also known as the black-headed bull ant, is a species of ant endemic to Australia. A member of the genus Myrmecia in the subfamily Myrmeciinae, it was first described by Austrian entomologist Gustav Mayr in 1862. These ants are large, varying from 19 to 23 millimetres (0.75 to 0.91 in) in length. However, colonies contain workers that are much smaller, usually half the size of normal workers. The queens are the largest while the males are the smallest, which can be easily identified due to their small mandibles.
Mainly nocturnal M. nigriceps is found in hot hilly areas and woodland, nesting underground in mounds. The ant's diet consists of sweet liquids from flowering plants and invertebrate prey, which are fed to the carnivorous larvae. Spiders are known to eat these ants, and echidnas consume the larvae and pupae. Like other Myrmecia species, M. nigriceps ants possess a powerful and painful sting, and the venom is capable of inducing allergic reactions in sensitive people. (Full article...)
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Theridion grallator, also known as the Hawaiian happy-face spider, is a spider in the familyTheridiidae that resides on the Hawaiian Islands. T. grallator gets its vernacular name of "Hawaiian happy-face spider" from the unique patterns superimposed on its abdomen, specifically those that resemble a smiley face. T. grallator is particularly notable because of its wide range of polymorphisms that may be studied to allow a better understanding of evolutionary mechanisms. In addition to the variety of color polymorphisms present, T. grallator demonstrates the interesting quality of diet-induced color change, in which its appearance temporarily changes as it metabolizes various food items. (Full article...)
Slimonidae (the name deriving from the type genus Slimonia, which is named in honor of Welsh fossil collector and surgeon Robert Slimon) is a family of eurypterids, an extinct group of aquatic arthropods. Slimonids were members of the superfamily Pterygotioidea and the family most closely related to the derived pterygotid eurypterids, which are famous for their cheliceral claws and great size. Many characteristics of the Slimonidae, such as their flattened and expanded telsons (the posteriormost division of their bodies), support a close relationship between the two groups.
Slimonids are defined as pterygotioid eurypterids with swimming legs similar to those of the type genus, Slimonia, and the second to fifth pair of appendages being non-spiniferous. The family contains only two genera, the almost completely known Slimonia and Salteropterus, which is known only from the telson and the metastoma (a large plate part of the abdomen).
Both slimonid genera preserve flattened and expanded telsons that end in elongated telson spikes. The discovery of several articulated specimens of Slimonia with the tail segments preserved in tight curves, suggesting that the tail segments were considerably more flexible than previously thought and would have been capable of considerable side-to-side movement. Unlike the related pterygotids, the slimonids did not possess robust and powerful cheliceral claws and as such, these telson spikes may have been the primary weaponry used by Slimonia, although this theory is considered unlikely by contemporary researchers. The telson spike of Salteropterus was likely not used as a weapon and was highly distinct and different from that of any other eurypterid. (Full article...)
Unlike the closely related wasps and ants, who are carnivorous/omnivorous, bees are herbivores that specifically feed on nectar (nectarivory) and pollen (palynivory), the former primarily as an carbohydrate source for metabolic energy, and the latter primarily for protein and other nutrients for their larvae. They are found on every continent except Antarctica, and in every habitat on the planet that contains insect-pollinatedflowering plants. The most common bees in the Northern Hemisphere are the Halictidae, or sweat bees, but they are small and often mistaken for wasps or flies. Bees range in size from tiny stingless bee species, whose workers are less than 2 millimeters (0.08 in) long, to the leafcutter bee Megachile pluto, the largest species of bee, whose females can attain a length of 39 millimeters (1.54 in). Vertebrate predators of bees include primates and birds such as bee-eaters; insect predators include beewolves and dragonflies.
Bees are best known to humans for their ecological roles as pollinators and, in the case of the best-known species, the western honey bee, for producing honey, a regurgitated viscous mixture of digested monosaccharides kept as food storage of the bee colony. Pollination management via bees is important both ecologically and agriculturally, and the decline in wild bee populations has increased the demand and value of domesticatedpollination by commercially managed hives of honey bees. The analysis of 353 wild bee and hoverfly species across Britain from 1980 to 2013 found the insects have been lost from a quarter of the places they inhabited in 1980. Human beekeeping or apiculture (meliponiculture for stingless bees) has been practiced as a discipline of animal husbandry for millennia, since at least the times of Ancient Egypt and Ancient Greece. Bees have appeared in mythology and folklore, through all phases of art and literature from ancient times to the present day, although primarily focused in the Northern Hemisphere where beekeeping is far more common. In Mesoamerica, the Mayans have practiced large-scale intensive meliponiculture since pre-Columbian times. (Full article...)
Myrmecophytes (/mərˈmɛkəfaɪt/; literally "ant-plant") are plants that live in a mutualistic association with a colony of ants. There are over 100 different genera of myrmecophytes. These plants possess structural adaptations in the form of domatia where ants can shelter, and food bodies and extrafloral nectaries that provide ants with food. In exchange for these resources, ants aid the myrmecophyte in pollination, seed dispersal, gathering of essential nutrients, and defense. Domatia adapted specifically to ants may be called myrmecodomatia. (Full article...)
H. socialis is the type species of Hughmilleriidae, a eurypterid family classified in the superfamily Pterygotioidea that is differentiated by their streamlined bodies, the enlargement of their medium-sized chelicerae and the presence of paired spines on the walking appendages. With the biggest specimen measuring 20 centimetres (8 inches) in length, Hughmilleria is considered a eurypterid of small size. (Full article...)
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Arthropleura sp., juvenile specimen from Montceau-les-Mines (France)
Arthropleura, from Ancient Greek ἄρθρον (árthron), meaning "joint", and πλευρά (pleurá), meaning "rib", is an extinct genus of massive myriapod that lived in what is now Europe and North America around 344 to 292 million years ago, from the Viséan stage of the lower Carboniferous period to the Sakmarian stage of the lower Permian period. It is a millipede, and was capable of reaching at least 2 metres (6 ft 7 in) in length, possibly up to over 2.5 metres (8 ft 2 in), making it the largest known land arthropod of all time. Arthropleura is known from body fossils as well as trace fossils, particularly giant trackways up to 50 centimetres (20 in) wide, and potentially also large burrows. It lived in open, sparsely wooded environments near water, and was possibly amphibious. (Full article...)
Compared to other stick insects (order Phasmatodea), the genus Timema is considered basal; that is, the earliest "branch" to diverge from the phylogenetic tree that includes all Phasmatodea. To emphasize this outgroup status, all stick insects not included in Timema are sometimes described as "Euphasmatodea."
Five of the twenty-one species of Timema are parthenogenetic, including two species that have not engaged in sexual reproduction for one million years, the longest known asexual period for any insect. (Full article...)
The following are images from various arthropod-related articles on Wikipedia.
Image 1This Zoea-stage larva is hardly recognisable as a crab, but each time it sheds its cuticle it remodels itself, eventually taking on its final crab form (from Arthropod exoskeleton)
Image 2 This fully-grown robber crab has tough fabric forming its joints, delicate biomineralized cuticle over its sensory antennae, optic-quality over its eyes, and strong, calcite-reinforced chitin armouring its body and legs; its pincers can break into coconuts (from Arthropod exoskeleton)
Image 10Mature queen of a termite colony, showing how the unsclerotised cuticle stretches between the dark sclerites that failed to stretch as the abdomen grew to accommodate her ovaries (from Arthropod exoskeleton)
Image 13The house centipedeScutigera coleoptrata has rigid sclerites on each body segment. Supple chitin holds the sclerites together and connects the segments flexibly. Similar chitin connects the joints in the legs. Sclerotised tubular leg segments house the leg muscles, their nerves and attachments, leaving room for the passage of blood to and from the hemocoel (from Arthropod exoskeleton)
Image 14 Honeybee larvae have flexible but delicate unsclerotised cuticles. (from Arthropod exoskeleton)
Image 15Formation of anterior segments across arthropod taxa based on gene expression and neuroanatomical observations, Note the chelicera(Ch) and chelifore(Chf) arose from somite 1 and thus correspond to the first antenna(An/An1) of other arthropods. (from Chelicerata)
Image 17Ghost crab, showing a variety of integument types in its exoskeleton, with transparent biomineralization over the eyes, strong biomineralization over the pincers, and tough chitin fabric in the joints and the bristles on the legs (from Arthropod exoskeleton)
Image 18Body structure of a typical crustacean – krill (from Crustacean)
Image 26In honeypot antrepletes, the abdomens of the workers that hold the sugar solution grow vastly, but only the unsclerotised cuticle can stretch, leaving the unstretched sclerites as dark islands on the clear abdomen (from Arthropod exoskeleton)
Image 32Some of the various hypotheses of myriapod phylogeny. Morphological studies (trees a and b) support a sister grouping of Diplopoda and Pauropoda, while studies of DNA or amino acid similarities suggest a variety of different relationships, including the relationship of Pauropoda and Symphyla in tree c. (from Myriapoda)
Image 33Reconstruction of Mollisonia plenovenatrix, the oldest known arthropod with confirmed chelicerae (from Chelicerata)
The Forest scorpion (Cercophonius squama) is a scorpion native to southeastern Australia and Tasmania. The body is 25 to 40 millimetres (0.98 to 1.57 in) long, and coloured creamy yellow to orange brown with dark brown variegations. The legs are yellow with some dark brown pigment.
The cryptic mantis (Sibylla pretiosa) is a mantid native to southern Africa. Its common name comes from its ability to grow asymmetrically to match the vegetation of its environment. They have unusual leaf-like projections on the joint of their four walking legs and leaf-like wings, generally the only green portion of the insect's otherwise brown and mottled exoskeleton. Adult females grow to 5–6 cm (2.0–2.4 in) in length while the males are generally about 1 cm smaller.
Gonepteryx rhamni, also known as the common brimstone, is a butterfly in the family Pieridae. It lives throughout the Palearctic zone and is commonly found across Europe, Asia and North Africa. The butterfly relies on two species of buckthorn as hosts for its eggs and larvae, which influences its geographic range and distribution as these plants are commonly found in wetlands. After spending the summer feeding, adults travel to woodland areas to spend seven months hibernating. In spring when their host plants have developed, they return to the wetlands to breed and lay eggs. Both the larval and adult forms of the species have protective coloration and behaviour that decreases their chances of being recognised and preyed upon. The adult common brimstone has sexual dichromism in its wing coloration and iridescence; the male (pictured) has yellow wings and iridescence, while females have greenish-white wings and are not iridescent.
The orb-weaver spiders (family Araneidae) are the familiar builders of spiral wheel-shaped webs often found in gardens, fields and forests. The family is a large one, including over 2800 species in over 160 genera worldwide, making it the third largest known (behind Salticidae and Linyphiidae). The web has always been thought of as an engineering marvel.
The monarch butterfly (Danaus plexippus) is one of the best-known species of butterfly. Its wings feature an easily recognizable orange and black pattern, with a wingspan of 8.9–10.2 centimetres (3½–4 in).
Papilio polymnestor, the blue Mormon, is a species of swallowtail butterfly found in southern India and Sri Lanka. It is a woodland species, often seen on forest paths and near streams. The larvae feed on trees in the family Rutaceae, such as citrus. Young larvae are green with white markings and position themselves on the upper surface of leaves, relying on their cryptic colouring, which resembles bird droppings, for protection. Older larvae seek less conspicuous locations, and have a unique habit of securing their balance by weaving silk on the substratum. This adult male P. polymnestor butterfly was photographed in the Indian state of Kerala.
Xylotrupes socrates (Siamese rhinoceros beetle, or "fighting beetle"), male, on a banana leaf. This scarab beetle is particularly known for its role in insect fighting in Northern Laos and Thailand.
The longtail tadpole shrimp (Triops longicaudatus) is a freshwater crustacean resembling a miniature horseshoe crab. It is one of the oldest animal species still in existence. Like its relative Triops cancriformis, the longtail tadpole shrimp is considered a living fossil because its basic prehistoric morphology has changed little in the last 70 million years, exactly matching ancient fossils.
A honey bee extracts nectar from a flower using its proboscis. Tiny hairs covering the bee's body maintain a slight electrostatic charge, causing pollen from the flower's anthers to stick to the bee's hairs, allowing for pollination when the bee moves on to another flower.
A fishing spider with two of its legs missing. Most likely some predator (a bird, or given its habitat a large fish or frog) grasped the spider by the missing limbs which were jettisoned by the spider in response, a process known as autotomy.
A macro view of a Gonia capitatafly feeding on honey, showing its proboscis and pedipalps (the two appendages protruding from the proboscis), two types of insect mouthparts. The proboscis actually comprises the labium, a quadrupedal structure, and a sponge-like labellum at the end. Flies eat solid food by secreting saliva and dabbing it over the food item. As the saliva dissolves the food, the solution is then drawn up into the mouth as a liquid. The labellum's surface is covered by minute food channels which form a tube leading to the esophagus, and food is drawn up the channels by capillary action.
The head and mandibles of an Australianbull ant. Insect mandibles grasp, crush, or cut the insect’s food, or defend against predators or rivals. These mandibles move in the horizontal plane unlike those of the vertebrates.
The light blue soldier crab (Mictyris longicarpus) inhabits beaches in the Indo-Pacific region. Soldier crabs filter sand or mud for microorganisms. They congregate during the low tide, and bury themselves in a corkscrew pattern during high tide, or whenever they are threatened.
Danaus genutia, the common tiger or striped tiger, is a species of brush-footed butterfly found in Sri Lanka, India, Myanmar, south-eastern Asia and Australia. It prefers areas of moderate to heavy rainfall, and typical habitats include scrubby jungle, deciduous forests and fallow land near habitations. The insect sequesters toxins from plants, and advertises its unpalatability by having prominent markings and striking colour patterns. This adult male common tiger, of the subspecies D. g. genutia, was photographed in Kerala, India.
The Ozyptila praticola species of crab spider is found throughout Europe and the Middle East. They do not build webs to trap prey, but are active hunters. Crab spiders are so named because of their first two pairs of legs, which are held out to the side giving them a crab-like appearance. Also, like crabs, these spiders move sideways and backwards more easily than forwards.
Sympetrum danae, the black darter or black meadowhawk, is a species of dragonfly found in northern Europe, Asia, and North America. Both sexes are black and yellow, but the abdomen of the male is largely black while that of the female is largely yellow. Breeding takes place in shallow acidic pools, lake margins and ditches in lowland heaths and moorland bogs. The female lays her eggs during flight by dipping the tip of her abdomen into the water. The eggs hatch the following spring, the larvae developing very rapidly and emerging as adults in as little as two months. The male seen here is perched on a frond of bracken on Warren Heath in Hampshire, England.
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