Goio-Erê Formation

Goio-Erê Formation
Goio-Erê Formation
Stratigraphic range: Aptian-Campanian?; ~125–71 Ma PreꞒ Ꞓ O S D C P T J K Pg N
Type	Geological formation
Unit of	Caiuá Group (Bauru Basin)
Underlies	Alluvium
Overlies	Unconformity with Lower Cretaceous rocks
Lithology
Primary	Sandstone
Location
Coordinates	23°48′S 53°06′W / 23.8°S 53.1°W
Approximate paleocoordinates	28°54′S 32°42′W / 28.9°S 32.7°W
Country	Brazil
Extent	Bauru Sub-basin, Paraná Basin
	Goio-Erê Formation (Brazil)

The Goio-Erê Formation is a geological formation in Brazil. It is sometimes thought to be deposited between the Turonian and Campanian stages of the Late Cretaceous,^[1] but an Aptian-Albian date has also been proposed.^[2]^[3] It primarily consists of sandstone and was deposited in a desert environment. It is known for its exceptional 3-D preservation of fossils at the Cemitério dos Pterossauros ("Pterosaur Graveyard") Quarry, located in the municipality of Cruzeiro do Oeste.^[2]^[3] Animals recovered from this site include the pterosaurs Keresdrakon, Caiuajara and Torukjara, the dinosaurs Berthasaura and Vespersaurus, and the iguanian lizard Gueragama. A few studies have alternatively assigned the Cemitério dos Pterossauros site to the younger Rio Paraná Formation, however.^[4]^[5]

Fossil content

Dinosaurs

Dinosaurs from the Goio-Erê Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Berthasaura	B. leopoldinae^[3]	Cemitério dos Pterossauros Quarry			A berthasaurid ceratosaur.
Vespersaurus	V. paranaensis^[4]	Cemitério dos Pterossauros Quarry			A noasaurid ceratosaur.

Pterosaurs

Pterosaurs from the Goio-Erê Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Caiuajara	C. dobruskii^[1]	Cemitério dos Pterossauros Quarry			A tapejarid pterosaur.
Keresdrakon	K. vilsoni^[2]	Cemitério dos Pterossauros Quarry			An azhdarchoid pterosaur.
Torukjara	T. bandeirae^[6]	Cemitério dos Pterossauros Quarry			A tapejarid pterosaur.

Squamates

Squamates from the Goio-Erê Formation
Genus	Species	Location	Stratigraphic position	Material	Notes	Images
Gueragama	G. sulamericana^[7]	Cemitério dos Pterossauros Quarry		Jaw fragments	An iguanian lizard.

References

^ ^a ^b Manzig, Paulo C.; Kellner, Alexander W.A.; Weinschütz, Luiz C.; Fragoso, Carlos E.; Vega, Cristina S.; Guimarães, Gilson B.; Godoy, Luiz C.; Liccardo, Antonio; Ricetti, João H.Z.; de Moura, Camila C. (2014-08-13). "Discovery of a Rare Pterosaur Bone Bed in a Cretaceous Desert with Insights on Ontogeny and Behavior of Flying Reptiles". PLOS ONE. 9 (8): e100005. Bibcode:2014PLoSO...9j0005M. doi:10.1371/journal.pone.0100005. ISSN 1932-6203. PMC 4131874. PMID 25118592.
^ ^a ^b ^c Kellner, Alexander W.A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renam A.M.; Sayão, Juliana M. (2019). "A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert". Anais da Academia Brasileira de Ciências. 91 (suppl 2): e20190768. doi:10.1590/0001-3765201920190768. ISSN 0001-3765. PMID 31432888.
^ ^a ^b ^c de Souza, Geovane Alves; Soares, Marina Bento; Weinschütz, Luiz Carlos; Wilner, Everton; Lopes, Ricardo Tadeu; de Araújo, Olga Maria Oliveira; Kellner, Alexander Wilhelm Armin (2021-11-18). "The first edentulous ceratosaur from South America". Scientific Reports. 11 (1): 22281. Bibcode:2021NatSR..1122281D. doi:10.1038/s41598-021-01312-4. ISSN 2045-2322. PMC 8602317. PMID 34795306.
^ ^a ^b Langer, Max Cardoso; de Oliveira Martins, Neurides; Manzig, Paulo César; de Souza Ferreira, Gabriel; de Almeida Marsola, Júlio César; Fortes, Edison; Lima, Rosana; Sant’ana, Lucas Cesar Frediani; da Silva Vidal, Luciano; da Silva Lorençato, Rosangela Honório; Ezcurra, Martín Daniel Ezcurra (2019). "A new desert-dwelling dinosaur (Theropoda, Noasaurinae) from the Cretaceous of south Brazil". Scientific Reports. 9 (1): 9379. Bibcode:2019NatSR...9.9379L. doi:10.1038/s41598-019-45306-9. PMC 6594977. PMID 31243312.
^ Ribeiro, Theo Baptista; Brito, Paulo Marques Machado; Gomes Da Costa Pereira, Paulo Victor Luiz (2024). "The predominance of teeth in the non-avian dinosaur record from Cretaceous Brazil: A review". Historical Biology. 36 (10): 2019–2034. Bibcode:2024HBio...36.2019R. doi:10.1080/08912963.2023.2238965.
^ Pêgas, Rodrigo V. (2024-06-10). "A taxonomic note on the tapejarid pterosaurs from the Pterosaur Graveyard site (Caiuá Group, ?Early Cretaceous of Southern Brazil): evidence for the presence of two species". Historical Biology: 1–22. doi:10.1080/08912963.2024.2355664. ISSN 0891-2963.
^ Simões, Tiago R.; Wilner, Everton; Caldwell, Michael W.; Weinschütz, Luiz C.; Kellner, Alexander W. A. (November 2015). "A stem acrodontan lizard in the Cretaceous of Brazil revises early lizard evolution in Gondwana". Nature Communications. 6 (1): 8149. Bibcode:2015NatCo...6.8149S. doi:10.1038/ncomms9149. ISSN 2041-1723. PMC 4560825. PMID 26306778.

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[manzig2014-1] Manzig, Paulo C.; Kellner, Alexander W.A.; Weinschütz, Luiz C.; Fragoso, Carlos E.; Vega, Cristina S.; Guimarães, Gilson B.; Godoy, Luiz C.; Liccardo, Antonio; Ricetti, João H.Z.; de Moura, Camila C. (2014-08-13). "Discovery of a Rare Pterosaur Bone Bed in a Cretaceous Desert with Insights on Ontogeny and Behavior of Flying Reptiles". PLOS ONE. 9 (8): e100005. Bibcode:2014PLoSO...9j0005M. doi:10.1371/journal.pone.0100005. ISSN 1932-6203. PMC 4131874. PMID 25118592.

[Kellner2019-2] Kellner, Alexander W.A.; Weinschütz, Luiz C.; Holgado, Borja; Bantim, Renam A.M.; Sayão, Juliana M. (2019). "A new toothless pterosaur (Pterodactyloidea) from Southern Brazil with insights into the paleoecology of a Cretaceous desert". Anais da Academia Brasileira de Ciências. 91 (suppl 2): e20190768. doi:10.1590/0001-3765201920190768. ISSN 0001-3765. PMID 31432888.

[Berthasaura-3] Souza, Geovane Alves; Soares, Marina Bento; Weinschütz, Luiz Carlos; Wilner, Everton; Lopes, Ricardo Tadeu; de Araújo, Olga Maria Oliveira; Kellner, Alexander Wilhelm Armin (2021-11-18). "The first edentulous ceratosaur from South America". Scientific Reports. 11 (1): 22281. Bibcode:2021NatSR..1122281D. doi:10.1038/s41598-021-01312-4. ISSN 2045-2322. PMC 8602317. PMID 34795306.

[langer2019-4] Langer, Max Cardoso; de Oliveira Martins, Neurides; Manzig, Paulo César; de Souza Ferreira, Gabriel; de Almeida Marsola, Júlio César; Fortes, Edison; Lima, Rosana; Sant’ana, Lucas Cesar Frediani; da Silva Vidal, Luciano; da Silva Lorençato, Rosangela Honório; Ezcurra, Martín Daniel Ezcurra (2019). "A new desert-dwelling dinosaur (Theropoda, Noasaurinae) from the Cretaceous of south Brazil". Scientific Reports. 9 (1): 9379. Bibcode:2019NatSR...9.9379L. doi:10.1038/s41598-019-45306-9. PMC 6594977. PMID 31243312.

[5] Ribeiro, Theo Baptista; Brito, Paulo Marques Machado; Gomes Da Costa Pereira, Paulo Victor Luiz (2024). "The predominance of teeth in the non-avian dinosaur record from Cretaceous Brazil: A review". Historical Biology. 36 (10): 2019–2034. Bibcode:2024HBio...36.2019R. doi:10.1080/08912963.2023.2238965.

[6] Pêgas, Rodrigo V. (2024-06-10). "A taxonomic note on the tapejarid pterosaurs from the Pterosaur Graveyard site (Caiuá Group, ?Early Cretaceous of Southern Brazil): evidence for the presence of two species". Historical Biology: 1–22. doi:10.1080/08912963.2024.2355664. ISSN 0891-2963.

[7] Simões, Tiago R.; Wilner, Everton; Caldwell, Michael W.; Weinschütz, Luiz C.; Kellner, Alexander W. A. (November 2015). "A stem acrodontan lizard in the Cretaceous of Brazil revises early lizard evolution in Gondwana". Nature Communications. 6 (1): 8149. Bibcode:2015NatCo...6.8149S. doi:10.1038/ncomms9149. ISSN 2041-1723. PMC 4560825. PMID 26306778.

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